Association of transcription marks by phylostratum. Log-odds of gene counts as enrichment. A) combination of three transcriptional hallmarks: CpG islands, H3K4me3 peaks and DNAseI sensitivity hotspots. B) to D) profiles for single transcription marks, separately for unidirectional promotors (black lines) and bidirectional promotors (red lines). B) CpG islands, C) H3K4me3 peaks, D) DNAseI sensitivity hotspots. * Hypergeometric test, FDR corrected, p<0.01.