Translocation of trnQ between the 12S rRNA ( rrnS ) and control region (CR) observed in a sea spider, Achelia bituberculata , two terrestrial spiders, Ornithictinus huwena and Habronattus oregonensis , and a collembolan, Tetrodontophora bielanensis. Compared to the typical arthropod trnI-trnQ-M arrangement between nad2 and the CR, the trnQ gene in A. bituberculata is found between rrnS and the CR. Such trnQ translocation appears in the two terrestrial spiders, O. huwena and H. oregonensis, and in a collembolan, T. bielanensis. Refer to the Fig. 1 legend for abbreviations of components. The Q position is highlighted with an asterisk (*) and a yellow box. Bold underlines indicate the reversed orientation of gene transcription.