Changes in gene order in mitochondrial genomes of Arachnida compared to the putative ancestral arthropod gene order. Transfer RNA genes are labelled according to the one letter amino acid code. Genes marked white show the same relative position as in the arthropod ground pattern; genes marked orange have relative positions differing from the arthropod ground pattern; the gene marked black indicates a duplicated rrnL gene in Leptotrombidium pallidum. Horizontal lines above genes illustrate adjacent genes which were probably translocated together; dotted lines indicate regions where tandem duplication and random deletion events may have occurred; connected arrows show adjacent genes which have switched their position, making it difficult to assess which gene was translocated. Braces accentuate the duplicated regions in the mitochondrial genome of Metaseiulus occidentalis. lnr: large non-coding region, putative mitochondrial control region; other non-coding regions (> 50 bp) are illustrated by gaps between genes. Numbers refer to rearrangement events, compare Fig. 6. For GenBank accession numbers see Table 2.