Table 4 Parameter estimates and likelihood ratio tests for the site models

M0 (one ratio model)

1

-47935.395

ω=0.143

-

-

-

None

 M1: Neutral (k=2)

1

-47271.650

p ₀ =0.774, (p ₁ =0.226)

-

-

-

Not allowed

 M2 : Selction (k=3)

3

-47271.650

p ₀ =0.774, p ₁ =0.048, (p ₂=0.178), ω₂=1.000

M2 vs M1: 0.000

1

1.000

None

 M3: discrete (K=5)

5

-46728.169

p ₀ =0.244, p ₁ =0.389, p ₂ =0.263, p ₃ =0.094, p ₄ =0.011, ω₀=0.010, ω₁=0.085, ω₂=0.317, ω₃=0.799, ω₄=6.007

M3 vs M0: 2414.452

4

0.000

1 (p>0.95)

 M7: beta

2

-46761.572

p=0.513, q=1.910

-

-

-

Not allowed

 M8: beta & w>1

4

-46732.126

p ₀ =0.947, p=0.589, q=2.892, (p ₁ =0.053) ω=1.293

M8 vs M7: 58.891

2

0.000

6 (p>0.95), 3 (p>0.95)