Cladogram for changes in gene order of lophotrochozoan taxa (only changes in protein-coding and rRNA genes were analysed). The translocation of a gene or a gene block is treated as an apomorphic feature (small box) with numbers according to translocated genes in the table below. "x" indicates derived gene positions, circles stand for an unvaried order. "(x)" symbolizes that although the position of the gene is now different there is evidence that it. Questionmarks indicate missing sequence data or putative secondary events complicating the interpretation. Changes not mentioned in the table: (10) translocation of cox3/nad3; (11) translocation of rrnS/rrnL/nad1; (12) translocation of nad6; (13) large inversion of a segment spanning from rrnS to nad5; (14) translocation of atp6. See text for further details.