Nucleosomal features that distinguish the four classes. Average values that correspond to mean histone acetylation levels  (H3K9, H3K14, and H4 acetylation), the turnover rate of H3 histone , nucleosome fuzziness , and the mean levels of the occupancy for the seven remodelers  are shown for the four classes. For properties except remodeler occupancy, we assigned each TFBS with the corresponding value of its nearest probe. For remodeler occupancy, we assigned each promoter with the corresponding value. Values in each property were normalized, such that their means are zero and standard deviations are one. Error bars were calculated by bootstrapping.