From: Inferring linkage disequilibrium from non-random samples†
MM
Mm
mm
AA
Aa
aa
Q 2
2Q(1 - Q)
(1 - Q)2
QR
Q + R - 2QR
(1 - Q)(1 - R)
R 2
2R(1 - R)
(1 - R)2
n 11
n 12
n 13
n 21
n 22
n 23
n 31
n 32
n 33