Metabolic features of C. variabile DSM 44702 relevant for the lifestyle on the surface of smear-ripened cheeses and during cheese ripening. The selected metabolic and regulatory features were deduced from the annotation of the complete genome sequence. Processes and relevant proteins associated with central carbohydrate metabolism, lipolysis and fatty acid metabolism, proteolysis and amino acid uptake, iron acquisition, stress responses, food quality, as well as flavor generation are indicated and labeled. Relevant transport systems are shown within the cell membrane. Numbers in parenthesis indicate the number of paralogs present in the C. variabile DSM 44702 genome. Relevant transcription regulators are shown by grey circles and boxed according to the physiological module they play a regulatory role in. Abbreviations: GABA, γ-amino butyric acid; PPP, pentose phosphate pathway; TCA, tricarboxylic acid cycle.